This book examines the profound evolutionary consequences of the Late Devonian extinctions and the various theories proposed to explain their occurrence. Only one group of four-limbed vertebrates exists on Earth, while other tetrapod-like fishes are extinct. This gap is why the idea of “fish with feet” seems so peculiar to us, yet such animals were once a vital part of our world. If the Devonian extinctions had not happened, members of these species, like the famous Acanthostega and Ichthyostega, might have continued to live in our rivers and lakes. Synthesizing decades of research and including a wealth of new discoveries, this accessible, comprehensive text explores the causes of the Devonian extinctions, the reasons vertebrates were so severely affected, and the potential evolution of the modern world if the extinctions had never taken place.